Laureato in Medicina presso l'Università di Napoli, con un Post dottorato: Patologia e Biologia Molecolare e cellulare presso la stessa Università. È Professore Ordinario di Patologia Generale presso il Dipartimento di Medicina, Chirurgia e Odontoiatria dell'Università degli Studi di Milano. È fondatore dell’Ifom (l'Istituto FIRC di Oncologia Molecolare) dove svolge la maggior parte delle sue ricerche ed è membro dell’Embo (European Molecular Biology Organization).
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